Allelopathic Assessment by Interaction Effect of Coconut Water (Cocos nucifera L.), and Dipping Time on Seed Germination of Four Cereal Seeds.

  • Yaseen A.R. Goran Department of Biology, College of Science ,Salahaddin University-Erbil, Kurdistan region-Iraq.
Keywords: Key words: allelopathy. barley, sorghum, wheat, soft, hard, coconut water, dipping time, concentration, interaction.


          Coconut water (Cocos nucifera L.) is a biological and sterile liquid from coconut kernel. It contains a variety of ionic materials, sugars, amino acids, and several plant growth regulators. This work has been achieved in plant physiology research laboratory in Biology Department, College of Science at Salahaddin University/Erbil during July to December 2019. The purpose of this study is to evaluate the effect of concentration, different dipping time and the interaction between both seed germination and seedling radicle and plumule length. The effect of different concentrations (1%, 2%, 3%, and 4% v/v) of coconut water and different dipping times (5, 10, 15, and 20 seconds) on four cereal seeds germination and seedling parameters, the seedling plumule and radicle lengths, also the interaction between both treatments were studied. In this study it claims that when using different concentrations, the best one is the 3% concentration that effects on percent germination of all seeds used in this study, where the higher germination percentage was recorded. The best effective concentration on radicle and plumule elongation was in 1% and all concentrations showed significant effection. The best active dipping time was at 5 seconds and others also showed significant effection on both radicles and plumules. In the case of interaction of both treatments the more positive effective interaction was at 5 seconds in 4% concentration on radicles and plumules.


Agampodi,V. A. and Bimali Jayawardena (2009). Effect of coconut (Cocos nucifera L.) water extracts on adventitious root development in vegetative propagation of Dracaena purplecompacta L. Acta Physiologiae Plantarum 2009, Volume 31, Issue 2, pp 279-284
Akae, T., Avci, M ., Dusunceli, F.(2004). Barely; post- harvest operations. OECD (2004). Consensus document on compositional considerations for new varieties of barely (Hordium vulgare); key food and feed nutrients and anti- nutrient s. Report No.12 Environment Directorate, OECD, Paris.
Akhiriana E., Samanhudi K., and Yunus A. (2019). Coconut Water and IAA Effect on the InVitro Growth of Tribulus terrestris L. Acta Universitatis Agriculturae et Silviculturae Mendelianae Brunensis, 67(1): 9 – 18.
Arditti, J. (2008). Micropropagation of Orchids, 2nd ed.; Blackwell Publishing: Oxford, UK; Volume II.
Asian and Pacific Coconut Community (APCC). International Codes and Standard for Aqueous Coconut Products, 2nd draft. Standards Task Force, Asian and Pacific Coconut Community: Jakarta, Indonesia, 1994.
Chan E and Elevitch C. R ,(2006). Cocos nusifera(coconut) tradiotional tree 2(1).
Chandra, G.S; proudlove, MO,; baxter E. D. The structure of barely endosperm; an important determinant of malt modification. J. Sci. Food Agric., 1999, 79, 37-46.
Chang-Hung, Chou (2006). “Introduction to allelopathy”, 2006, Part 1, 1-9.
Cholan, P.K.; Manipal, S.; Rajmohan, M.D.S.; Kumar, R.(2017)Assessment of the Anti-Fungal Activity of Green CW (Cocos nucifera l.) on Candida albicans An in-vitro study J. Pharm. Sci. & Res. V.ol. 9(2), 2017, 251-254
Dhillon B. S., Saxena S., Agrwal A and Tyagi R. K. (2006) Food grain crops in plant genetic resources. Narosa publishing house, New Delhi.
Ever, T; Millar, S. cereal grain structure and development: some implication for quality. J. cereal sci. 2002,36, 261 -284
Fincher, G. B.; Stone, B. A. Cell walls and their component in cereal grain technology. Adv. Cereal Sci. Technol. 1986, 8, 207-295.
Fraenkel, Gottfried S. (1959). “The raison d’Etre of secondary plant substances”, Science, 129 (3361).
Gamlath, J.; Aldred, G. P.; Panozzo, J. F. (2008). Barley (1-3; 1-4)- B-glucan and arabinoxylan content are related to kernel hardnes and water uptake. J. cereal Sci., 47, 365-371.
George, E.F.; Sherrington, P.D. (1984). Plant Propagation by Tissue Culture-Handbook and Directory of Commercial Laboratories; Exegetics Ltd: Edington, UK, 1984.
ICAR (2006) Hand book of Indian Agriculture. ICAR publication, New Delhi
Izydorczyk, M.S.; Dexter, J. E.; Desjardins, R. G.; Rossnagel, B.G.; Lagasse, S. L.; Hatcher, D. W. Roller milling condian hull-less barely; optimization of roller milling conditions and composition of mill streams. Cereal chem. 2003, 80, 637-644.
Jothi V. and Loomba S. (2013). Cocos nusifera; Its properties and contribution to densty; International Jornal Scientific study, volume 1. issue 3.
Kende, H.; Zeevaart. J. (1997). The five “Classical” plant hormones. Plant Cell, 9, 1197–1210.
Kobayashi, H.; Morisaki, N.; Tago, Y.; Hashimoto, Y.; Iwasaki, S.; Kawachi, E.; Nagata, R.; Shudo, K. (1997). Structural identification of a major cytokinin in coconut milk as 14-O-(3-O-[β-Dgalactopyranosyl-(1-->2)-α-D-galactopyranosyl-(1-->3)-α-L-arabinofuranosyl]-4-O-(α-L-arabinofuranosyl)-β-D-galactopyranosyl)-trans-zeatin riboside. Chem. Pharm. Bull., 45, 260–264.
Komgrit I., Suhaimine C. and Sompong T. (2011). Cytokinins and coconut water promoted abnormalities in zygotic embryo culture of oil palm Songklanakarin J. Sci. Technol. 33 (6), 653-657.
Narsih, Y. and Harijono Y. (2012). The study of germination and soaking time to improve nutritional quality of sorghum seed. International Food Research Journal 19(4): 1429-1432 Journal homepage:
Ogatis, R.A. (2015). A Comparative Evaluation of Coconut Water as Root Setting Medium for Rhizopora Stylosa Hypocotyl Propagation International Journal of Science and Research (IJSR) ISSN (Online): 2319-7064 Index Copernicus Value (2015): 78.96 | Impact Factor (2015): 6.391
Ram H. H. and Singh H. G. (2003) Crop breeding and genetics. Kalyani publication, new Delhi.
Rasmusson D. C. (1985). Barely. Crop science society of America publisher S.,Madison.
Rattan, S.I.S.; Clark, B.F.C. Kinetin delays the onset of ageing characteristics in human fibroblasts. Biochem. Biophys. Res. Commun. 1994, 201, 665–672.
Rice, E. L. (1984). Allelopathy, (1st Ed., November 1974 by the same editor) (2nd Ed.), Academic Press, pp. 422 p. ISBN 978-0-12-587058-0
Roger M., Reigosa J., Reigosa M., Pedrol J., Nuria J., and González L. (2006). Allelopathy: a physiological process with ecological implications, Springer, p. 1, ISBN 1- 4020-4279-5
Sandhya V.G., and Rajamohan T. (2008) Comparative evaluation of the hypolipidemic effects of coconut water and lovastatin in rats fed fat-cholesterol enriched diet. Food Chem. Toxicol., 45, 3585–3592.
Santos U., Kubo K., Ota T., Tadokoro T., and Maekawa A. (1996) Nutrient composition of coconuts water (Cocos nucifera L.). Food Chem., 57, 299–304.
Seow C.C., and Gwee C.N. (1997) Coconut milk: Chemistry and technology. Int. J. Food Sci. Tech., 32, 189–201.
Shewry, P. R. (1992) Barley seed storage proteins structure, synthesis, and deposition. In nitrogen metabolism of plant; pilbeam, D. J., Mengel, K., Eds.; Oxford university press; Oxford, Uk; pp 201-227.
United States Department of Agriculture (USDA). National Nutrient Database for Standard Reference, 2008. Nuts, coconut water [Online]. Available: foodcomp/cgi-bin/, accessed on 9 December 2009.
Vermeulen K., Strnad M., Kryštof V., Havlicek L., Van der Aa A., Lenjou M., Njis G., Rodrigus I., Stockman B., Van Onckelen H., Van Bockstaele D.R., and Berneman Z.N. (2002). Antiproliferative effect of plant cytokinin analogues with an inhibitory activity on cyclindependent kinases. Leukemia, 16, 299–305.
Willis, R. J. (2007). “The History of Allelopathy”, Springer: 3, ISBN 1-4020-4092-X, retrieved 2009-08-12.
Wood P. (2007) Cereal B –Glucan in diet and health J. Cereal Sci., 46, 230 -238.
How to Cite
Goran, Y. (2020) “Allelopathic Assessment by Interaction Effect of Coconut Water (Cocos nucifera L.), and Dipping Time on Seed Germination of Four Cereal Seeds.”, Zanco Journal of Pure and Applied Sciences, 32(4), pp. 122-134. doi: 10.21271/ZJPAS.32.4.15.
Biology and Medical Researches